Alex, Marty, Gloria and Melman are still fighting to get home to their beloved Big Apple. Their journey takes them through Europe where they find the perfect cover: a traveling circus, which they reinvent - Madagascar style.
Super villain Megamind's dreams have come true when he conquers the city's protector Metro Man gaining control of Metro City. But when a new villain is created and chaos runs rampant, the world's biggest "mind" might actually save the day.
In this story that occurs before his meeting with Shrek and his friends, the suave, swashbuckling Puss in Boots teams up with Humpty Dumpty and street-savvy Kitty Softpaws to steal the Goose that Lays the Golden Eggs.